Abdu, R., Tancredi, S., Abrahamson, D., & Balasubramaniam, R. (2025). Demonstrating mathematics learning as the emergence of eye–hand dynamic equilibrium. Educational Studies in Mathematics, 118(3), 505-528.
Agostinho, S., Tindall-Ford, S., Ginns, P., Howard, S. J., Leahy, W., and Paas, F. (2015). Giving learning a helping hand: finger tracing of temperature graphs on an iPad. Educ. Psychol. Rev. 27, 427–443. doi: 10.1007/s10648-015-9315-
Amin, T. G., & Haglund, J. (2015). Conceptual Metaphor and Embodied Cognition in Science Learning: Introduction to special issue. International Journal of Science Education. http://dx.doi.org/10.1080/09500693.2015.1025245
Babaei, B. (2020). Yādgiri-ye badanmand: Naqd-e ruykard-e asab-pazhuhi-ye falsafi be yādgiri va āmuzesh [Embodied learning: A critique of the philosophical neuroscientific approach to learning and education] [Doctoral dissertation, Ferdowsi University of Mashhad, Faculty of Education and Psychology]. https://ganj.irandoc.ac.ir
Bach, P. (2012). The embodiment of linguistic meaning: Key findings from psychology and neuroscience. Practical theories and empirical practice: A linguistic perspective, 35-54.
Ben-Ami, S., Buaron, B., Yaron, O., Keane, K., Sun, V. H., Phillips, F., ... & Mukamel, R. (2025). What the visual system can learn from the non-dominant hand: The effect of graphomotor engagement on visual discrimination. Memory & cognition, 53(1), 325-340. https://doi.org/10.3758/s13421-024-01628-2
Bilda, Z., Candy, L., & Edmonds, E. (2007). An embodied cognition framework for interactive experience. CoDesign, 3(2), 123-137. DOI: 10.1080/15710880701251443
Brennan, J. (2016). Naturalistic sentence comprehension in the brain. Language and Linguistics Compass, 10(7), 299-313. doi: 10.1111/lnc3.12198
Brown, W. S., & Reimer, K. S. (2013). EMBODIED COGNITION, CHARACTER FORMATION, AND VIRTUE: with Fraser Watts,“Embodied Cognition and Religion”; John A. Teske,“From Embodied to Extended Cognition”; Daniel H. Weiss,“Embodied Cognition in Classical Rabbinic Literature”; Léon Turner,“Individuality in Theological Anthropology and Theories of Embodied Cognition”; and Warren S. Brown and Kevin S. Reimer,“Embodied Cognition, Character Formation, and Virtue.”. Zygon®, 48(3), 832-845. https://doi.org/10.1111/zygo.12025
Burte, H., Gardony, A. L., Hutton, A., & Taylor, H. A. (2017). Think3d!: Improving mathematics learning through embodied spatial training. Cognitive Research: Principles and Implications, 2(1), 13. DOI 10.1186/s41235-017-0052-9
Butz, M. V. (2016). Toward a unified sub-symbolic computational theory of cognition. Frontiers in psychology, 7, 925. doi: 10.3389/fpsyg.2016.00925
Cappello, N., & Ortiz, D. C. (2025). Creative embodied learning about different identities. Thinking Skills and Creativity, 57, 101833. https://doi.org/10.1016/j.tsc.2025.101833
Castro-Alonso, J. C., Ayres, P., and Paas, F. (2014). Learning from observing hands in static and animated versions of non-manipulative tasks. Learn. Instr. 34, 11–21. doi:10.1016/j.learninstruc.2014.07.005
Castro-Alonso, J. C., Ayres, P., Zhang, S., De Koning, B. B., & Paas, F. (2024). Research avenues supporting embodied cognition in learning and instruction. Educational Psychology Review, 36(1), 10. https://doi.org/10.1007/s10648-024-09847-4
Ceja Salgado, O. (2022). Embodied learning in immersive virtual reality: The role of sensory-motor affordances in shaping conceptual and procedural understandings (Doctoral dissertation, UCL (University College London)).
Chandler, P., & Tricot, A. (2015). Mind your body: The essential role of body movements in children’s learning. Educational Psychology Review, 27(3), 365-370. DOI 10.1007/s10648-015-9333-3
Chang, C.-Y., Lin, H.-C., Yin, C., and Yang, K.-H. (2025). Generative AI-assisted reflective writing for improving students’ higher order thinking. Educ. Technol. Soc. 28, 270–285. doi: 0.30191/ETS.202501_28(1).TP03
Chang, W., & Aberash, A. (2026). Embodied learning in the digital age: assessing the impact of gesture-based educational technology on working memory capacity, spatial reasoning, and engagement. BMC psychology, 14(1), 331.https://doi.org/10.1186/s40359-026-04089-1
Chappell, K. (2024). Flowing with embodiment and materiality: Touch and time for new educational futures. In Creative ruptions for emergent educational futures (pp. 27-49). Cham: Springer Nature Switzerland.
Chen, C.-J., and Liu, P.-L. (2025). Combining immersive virtual reality with CLIL and TPR to enhance English as foreign language learners’ language acquisition. Educ. Technol. Soc. 28, 383–396. doi: 10.30191/ETS.202501_28(1).SP05
Chettaoui, N., Atia, A., and Bouhlel, M. S. (2022). Exploring the impact of interaction modality on students’ learning performance. J. Educ. Comput. Res. 60, 4–27. doi: 10.1177/07356331211027297
Claire, V. (2025). Developing enhanced embodied cognition using a sensory learning approach. Mental Health and Social Inclusion, 29(6), 669-671.
Clare, Y. (2018). Stanislavsky’s system as an enactive guide to embodied cognition?. In Embodied Cognition, Acting and Performance (pp. 43-63). Routledge. DOI: 10.1080/09540091.2016.1271397
Clifton, P. G., Chang, J. S. K., Yeboah, G., Doucette, A., Chandrasekharan, S., Nitsche, M., ... & Mazalek, A. (2016). Design of embodied interfaces for engaging spatial cognition. Cognitive Research: Principles and Implications 1, 1: 24. DOI 10.1186/s41235-016-0032-5
Conley, Q., Atkinson, R. K., Nguyen, F., and Nelson, B. C. (2020). Mantarayar:leveraging augmented reality to teach probability and sampling. Comput. Educ.153:103895. doi: 10.1016/j.compedu.2020.103895
Damsgaard, L., Nielsen, A. M. V., Gejl, A. K., Malling, A. S. B., Jensen, S. K., & Wienecke, J. (2022). Effects of 8 weeks with embodied learning on 5–6-year-old Danish children’s pre-reading skills and word reading skills: The PLAYMORE project, DK. Educational Psychology Review, 34(3), 1709-1737.
Damsgaard, L., Nielsen, A. M. V., Topor, M. K., Hansen, R. A., Jensen, S. K., Markers, R. L., ... & Wienecke, J. (2023). Embodied learning activities focusing on letter-sound knowledge increase spelling performance in 1st grade children with low and high reading ability. Educational Psychology Review, 35(3), 74.
Darejeh, A., Marcus, N., and Sweller, J. (2022). Increasing learner interactions with E-learning systems can either decrease or increase cognitive load depending on the nature of the interaction. Annee Psychol. 122, 405–437. doi: 10.3917/anpsy1.223.0405
Davis, J. I., Benforado, A., Esrock, E., Turner, A., Dalton, R. C., van Noorden, L., & Leman, M. (2012). Four applications of embodied cognition. Topics in cognitive science, 4(4), 786-793. DOI: 10.1111/j.1756-8765.2012.01225.x
De Koning, B. B., & Tabbers, H. K. (2011). Facilitating understanding of movements in dynamic visualizations: An embodied perspective. Educational Psychology Review, 23(4), 501-521. DOI 10.1007/s10648-011-9173-8
De Koning, B., Zhang, S., & Sepp, S. (2025). Integrating human movement in learning: advancements in language instruction, multimedia, and theory. Educational Psychology Review, 37(2), 51.
De Nooijer, J. A., Van Gog, T., Paas, F., and Zwaan, R. A. (2013). Eects of imitating gestures during encoding or during retrieval of novel verbs on children’s test performance. Acta Psychol. 144, 173–179. doi: 10.1016/j.actpsy.2013.05.013
DeSutter, D., & Stieff, M. (2017). Teaching students to think spatially through embodied actions: Design principles for learning environments in science, technology, engineering, and mathematics. Cognitive research: principles and implications, 2(1), 22. DOI 10.1186/s41235-016-0039-y
Dickinson, D., & Yates, M. (2026). Grounding verbs in action: The facilitative effect of potential physical interactions with verbs. Psychonomic Bulletin & Review, 33(1), 34.
Duijzer, C., Van den Heuvel-Panhuizen, M., Veldhuis, M., Doorman, M., & Leseman, P. (2019). Embodied learning environments for graphing motion: A systematic literature review. Educational Psychology Review, 31(3), 597-629https://doi.org/10.1007/s10648-019-09471-7
Farina, M. (2020). Embodied cognition: Dimensions, domains and applications. Adaptive Behavior, 28(5), 369–384. https://doi.org/10.1177/1059712320912963
Feiler, J. B. (2023). Do Embodied Representations Facilitate Mental Animation of Scientific Spatial Concepts? A Question of “STEMbodiment”. The University of Alabama.
Ferreira, J. M. (2021). What if we look at the body? An embodied perspective of collaborative learning. Educational Psychology Review, 33(4), 1455-1473. https://doi.org/10.1007/s10648-021-09607-8
Ferreira, J. M., & Ineson, G. (2026). Embodied learning in teacher education: Investigating student-teachers' experiences in engaging with embodied cognition theories. Teaching and Teacher Education, 171, 105335. https://doi.org/10.1016/j.tate.2025.105335
Fonteles, J. H., Cohn, C., Ayalon, E., Zhou, M., TS, A., Davalos, E., ... & Biswas, G. (2026). Analyzing embodied learning in classroom settings: A human-in-the-loop AI approach for multimodal learning analytics. Learning and Instruction, 103, 102274. https://doi.org/10.1016/j.learninstruc.2025.102274
Foolen, A. (2017). The hand in figurative thought and language. Angeliki Athanasiadou (Hg.), Studies in figurative thought and language, 179-198.
Friederici, A. D. (2011). The brain basis of language processing: from structure to function. Physiological reviews. doi:10.1152/physrev.00006.2011
Fu, R., & He, H. (2025). The influence of embodied cognition on the perception and representation of numbers: A meta-analysis. Acta Psychologica, 258, 105180. https://doi.org/10.1016/j.actpsy.2025.105180
Fugate, J. M., Macrine, S. L., & Cipriano, C. (2019). The role of embodied cognition for transforming learning. International Journal of School & Educational Psychology, 7(4), 274-288. DOI: 10.1080/21683603.2018.1443856
Gholami, T., & Hosseini Sarvari, H. (2025). Gozar az shenakht-gerayi-ye klasik ta edrak-e E4: Barrasi-yetahavvolat-e nazari dar enqelab-e sevom-e olum-e shenakhti [From classical cognitivism to E4 perception: Examining theoretical developments in the third revolution of cognitive science]. Pazhuhesh-haye Falsafi, 19(52), 215–240. https://doi.org/10.22034/jpiut.2025.69426.4239
Ginns, P., and King, V. (2021). Pointing and tracing enhance computer-based learning. Educ. Technol. Res. Dev. 69, 1387–1403. doi: 10.1007/s11423-021-099970
Ginns, P., Hu, F. T., Byrne, E., and Bobis, J. (2016). Learning by tracing worked examples. Appl. Cogn. Psychol. 30, 160–169. doi: 10.1002/acp.3171
Glenberg, A. M., Goldberg, A. B., & Zhu, X. (2011). Improving early reading comprehension using embodied CAI. Instructional Science, 39(1), 27-39. DOI 10.1007/s11251-009-9096-7
Hauk, O. (2016). Only time will tell–why temporal information is essential for our neuroscientific understanding of semantics. Psychonomic Bulletin & Review, 23(4), 1072-1079. DOI 10.3758/s13423-015-0873-9
Henriksen, A. H., Topor, M., Petersen, G. B., Rohde, R. A., Damsgaard, L., Nielsen, A. M. V., ... & Wienecke, J. (2025). Embodied Learning Using Virtual Reality and Mirrors Can Enhance Letter‐Sound Knowledge in 6–7‐Year‐Old Children. Journal of Computer Assisted Learning, 41(4), e70090. DOI:10.1111/jcal.70090
Holý, J., Bolek, V., & Müller, M. (2026). Working bodies, thinking minds: embodied cognition and attention in hybrid project work. International Journal of Organizational Analysis, 34(12), 104-125. https://doi.org/10.1108/IJOA-10-2025-6092
Hong, J. C., Tai, K. H., and Tai, T. Y. (2024). Enhancing mandarin learning through classroom response apps: a comparison of cognitive and aective factors in touch and shaking responses. J. Comput. Assist. Learn. 40, 3360–3368. doi: 10.1111/jcal.13066
Hotton, S., & Yoshimi, J. (2011). Extending dynamical systems theory to model embodied cognition. Cognitive Science, 35(3), 444-479.
Hsiao, H. S., Chang, I. H., Chen, Y. H., & Chen, J. C. (2025). Using gesture recognition with the memory strategy to improve preschoolers’ learning performance, motor skills, and executive function. Educational technology research and development, 73(3), 1791-1809. https://doi.org/10.1007/s11423-025-10471-4
Hsiao, H.-S., and Chen, J.-C. (2016). Using a gesture interactive game-based learning approach to improve preschool children’s learning performance and motor skills. Comput. Educ. 95, 151–162. doi: 10.1016/j.compedu.2016.01.005
Hsiao, H.-s., Chen, J.-c., Lin, C.-Y., and Chen, W.-N. (2018). e inuence of a gesture-based learning approach on preschoolers’ learning performance, motor skills, and motion behaviors. Interact. Learn. Environ. 26, 869–881. doi: 10.1080/10494820.2017.1419498
Hu, C., Chen, S., & Liu, Y. (2025). The Embodied Simulation of L2 Grammatical Aspect: Proficiency-Dependent Evidence from the Action-Sentence Compatibility Effect. Behavioral Sciences, 15(11), 1581. DOI: 10.3390/bs15111581
Hu, F.-T., Ginns, P., and Bobis, J. (2015). Getting the point: tracing worked examples enhances learning. Learn. Instr. 35, 85–93. doi: 10.1016/j.learninstruc.2014.10.002
Huang, Y., Richter, E., Kleickmann, T., and Richter, D. (2022). Class size afects preservice teachers’ physiological and psychological stress reactions: an experiment in a virtual reality classroom. Comput. Educ. 184:104503. doi: 10.1016/j.compedu.2022.104503
Humburg, M., Danish, J. A., Tu, X., Georgen, C., Carter, B., & Enyedy, N. (2026). Using scientific annotation tools to support collaborative embodied learning in elementary school classrooms. Journal of Science Education and Technology, 35(1), 261-282. https://doi.org/10.1007/s10956-025-10248-7
Hung, I.-C., and Chen, N.-S. (2018). Embodied interactive video lectures for improving learning comprehension and retention. Comput. Educ. 117, 116–131. doi: 10.1016/j.compedu.2017.10.005
Hung, I.-C., Lin, L.-I., Fang, W.-C., and Chen, N.-S. (2014). Learning with the body: an embodiment-based learning strategy enhances performance of comprehending fundamental optics. Interact. Comput. 26, 360–371. doi: 10.1093/iwc/iwu011
Ionescu, T., & Ilie, A. (2018). Language learning in preschool children: An embodied learning account. Early child development and care, 188(1), 4-15. DOI: 10.1080/03004430.2016.1189419
Jiang, H., & Wu, W. (2025). How embodied cognition affects vocal performance: an integrative review. Journal of Voice. https://doi.org/10.1016/j.jvoice.2025.11.005
Johnson-Glenberg, M. C., & Megowan-Romanowicz, C. (2017). Embodied science and mixed reality: How gesture and motion capture affect physics education. Cognitive research: principles and implications, 2(1), 24.
Johnson-Glenberg, M. C., and Megowan-Romanowicz, C. (2017). Embodied science and mixed reality: how gesture and motion capture afect physics education. Cognitive Research 2, 24–28. doi: 10.1186/s41235-017-0060-9
Johnson-Glenberg, M. C., Birchfeld, D. A., Tolentino, L., and Koziupa, T. (2014b). Collaborative embodied learning in mixed reality motion-capture environments: two science studies. J. Educ. Psychol. 106, 86–104. doi: 10.1037/a0034008
Johnson-Glenberg, M. C., Ly, V., Su, M., Zavala, R. N., Bartolomeo, H., and Kalina, E.(2020). Embodied agentic STEM education: eects of 3D VR compared to 2D PC. 2020 6th international conference of the immersive learning research network (iLRN), IEEE. 24–30. doi: 10.23919/iLRN47897.2020.9155155
Johnson-Glenberg, M. C., Savio-Ramos, C., and Henry, H. (2014a). “Alien health”: a nutrition instruction exergame using the Kinect sensor. Games Health J. 3, 241–251. doi: 10.1089/g4h.2013.0094
Kalenine, S., Ott, L., & Casalis, S. (2025). Improving lexico-semantic integration with gesture-enriched pictures: A word-learning study using the Picture-Word Interference paradigm. Memory & Cognition, 53(7), 2111-2125. DOI: 10.3758/s13421-025-01701-4
Kirschner, P. A. (2002). Cognitive load theory: Implications of cognitive load theory on the design of learning. Learning and instruction, 12(1), 1-10. www.elsevier.com/locate/learninstruc
Kontra, C., Goldin‐Meadow, S., & Beilock, S. L. (2012). Embodied learning across the life span. Topics in cognitive science, 4(4), 731-739. DOI: 10.1111/j.1756-8765.2012.01221.x
Korbach, A., Ginns, P., Brünken, R., and Park, B. (2020). Should learners use their hands for learning? Results from an eye-tracking study. J. Comput. Assist. Learn. 36, 102–113. doi: 10.1111/jcal.12396
Kuo, F.-R., Hsu, C.-C., Fang, W.-C., and Chen, N.-S. (2014). the efects of embodiment-based TPR approach on student English vocabulary learning achievement, retention and acceptance. J. King Saud Univ. Comput. Inf. Sci. 26, 63–70. doi: 10.1016/j.jksuci.2013.10.003
Kwon, K., Kim, H., Kim, K., Seo, M., and Brush, T. A. (2025). Computational thinking through embodied learning: a comparison of mixed-reality and unplugged activities. J. Res. Technol. Educ., 1–17. doi: 10.1080/15391523.2025.2492604
Lai, C.-J. (2024). Examining the impact of multimodal task design on English oral communicative competence in fourth-grade content-language integrated social studies: a quasi-experimental study. Asia-Pac. J. Second Foreign Lang. Educ. 9:64. doi: 10.1186/s40862-024-00289-7
Lajevardi, N., Narang, N. S., Marcus, N., and Ayres, P. (2017). Can mimicking gestures facilitate learning from instructional animations and static graphics? Comput. Educ. 110, 64–76. doi: 10.1016/j.compedu.2017.03.010
Lan, Y.-J., Fang, W.-C., Hsiao, I. Y., and Chen, N.-S. (2018). Real body versus 3D avatar:the efects of diferent embodied learning types on EFL listening comprehension. Educ. Technol. Res. Dev. 66, 709–731. doi: 10.1007/s11423-018-9569-y
Lazovic, M. (2025). Spatial resources in pre-service teachers’ instructional practices in VR tandems: co-constructing shared spaces and embodied spatial scaffolding. Frontiers in Communication, 10, 1519165.
Lebert, A., & Vilarroya, O. (2024). The links between experiential learning and 4E cognition. Annals of the New York Academy of Sciences, 1541(1), 37–52. https://doi.org/10.1111/nyas.15238
Lindgren, R., & Johnson-Glenberg, M. (2013). Emboldened by embodiment: Six precepts for research on embodied learning and mixed reality. Educational researcher, 42(8), 445-452. DOI: 10.3102/0013189X13511661
Lindgren, R., Tscholl, M., Wang, S., and Johnson, E. (2016). Enhancing learning and engagement through embodied interaction within a mixed reality simulation. Comput. Educ. 95, 174–187. doi: 10.1016/j.compedu.2016.01.001
Liu, S., Toreini, P., & Maedche, A. (2025). Mixed Reality Learning Systems with Head-Mounted Displays in Higher Education: A Systematic Review. Technology, Knowledge and Learning, 1-35.
Liyanawatta, M., Yang, S. H., Liu, Y. T., Zhuang, Y., and Chen, G. d. (2022). Audience participation digital drama-based learning activities for situational learning in the classroom. Br. J. Educ. Technol. 53, 189–206. doi: 10.1111/bjet.13160
Lorrimar, V. (2019). MIND UPLOADING AND EMBODIED COGNITION: A THEOLOGICAL RESPONSE: with Robert M. Geraci and Simon Robinson,“Introduction to the Symposium on Artificial Intelligence and Apocalypticism”; Beth Singler,“Existential Hope and Existential Despair in AI Apocalypticism and Transhumanism”; Michael Morelli,“The Athenian Altar and the Amazonian Chatbot: A Pauline Reading of Artificial Intelligence and Apocalyptic Ends”; Victoria Lorrimar,“Mind Uploading and Embodied Cognition: A Theological Response”; and Syed .... Zygon®, 54(1), 191-206. https://doi.org/10.1111/zygo.1
Lui, M., McEwen, R., and Mullally, M. (2020). Immersive virtual reality for supporting complex scientific knowledge: augmenting our understanding with physiological monitoring. Br. J. Educ. Technol. 51, 2181–2199. doi: 10.1111/bjet.13022
Lyu, C., & Deng, S. (2024). Effectiveness of embodied learning on learning performance: A meta-analysis based on the cognitive load theory perspective. Learning and Individual Differences, 116, 102564. https://doi.org/10.1016/j.lindif.2024.102564
Maalej, Z., & Yu, N. (2011). Introduction: Embodiment via body parts. In Embodiment via body parts: Studies from various languages and cultures (pp. 1-20). John Benjamins Publishing Company. DOI: 10.1080/10926488.2012.691771
Macken, L., and Ginns, P. (2014). Pointing and tracing gestures may enhance anatomy and physiology learning. Med. Teach. 36, 596–601. doi: 10.3109/0142159X.2014.899684
Makransky, G., & Petersen, G. B. (2021). The cognitive affective model of immersive learning (CAMIL): A theoretical research-based model of learning in immersive virtual reality. Educational psychology review, 33(3), 937-958. https://doi.org/10.1007/s10648-020-09586-2
Malinverni, L., Schaper, M.-M., and Pares, N. (2016). An evaluation-driven design approach to develop learning environments based on full-body interaction. Educ. Technol. Res. Dev. 64, 1337–1360. doi: 10.1007/s11423-016-9468-z
Manches, A., Nygren, M., Price, S., & Revueltas Roux, A. (2025). Challenges of translational education research: Teachers’ interpretations of embodied learning. Cogent Education, 12(1), 2490443. https://doi.org/10.1080/2331186X.2025.2490443
Mancusi, F., Neumann, W. P., Pierri, F., & Fruggiero, F. (2025). The Embodied Cognition paradigm: a novel approach to advancing Human-Robot Collaboration research. IFAC-PapersOnLine, 59(10), 2646-2651. https://doi.org/10.1016/j.ifacol.2025.09.445.
Mansour, N., Aras, C., Staarman, J. K., & Alotaibi, S. B. M. (2025). Embodied learning of science concepts through augmented reality technology. Education and Information Technologies, 30(6), 8245-8275.
Mayer, R. E., and DaPra, C. S. (2012). An embodiment efect in computer-based learning with animated pedagogical agents. J. Exp. Psychol. Appl. 18, 239–252. doi: 10.1037/a0028616
McCaffrey, G. (2024). Enactivism: Embodied cognition, sense‐making, and nursing. Nursing Inquiry, 31(4), e12672. DOI: 10.1111/nin.12672
Mierowsky, R., Marcus, N., and Ayres, P. (2020). Using mimicking gestures to improve observational learning from instructional videos. Educ. Psychol. 40, 550–569. doi: 10.1080/01443410.2019.1650896
Moreau, D., Morrison, A. B., & Conway, A. R. (2015). An ecological approach to cognitive enhancement: Complex motor training. Acta psychologica, 157, 44-55.
Nathan, M. J., & Swart, M. I. (2021). Materialist epistemology lends design wings: Educational design as an embodied process. Educational Technology Research and Development, 69(4), 1925-1954. https://doi.org/10.1007/s11423-020-09856-4
Nathan, M. J., & Walkington, C. (2017). Grounded and embodied mathematical cognition: Promoting mathematical insight and proof using action and language. Cognitive research: principles and implications, 2(1), 9. DOI 10.1186/s41235-016-0040-5
Nguyen, D. J., & Larson, J. B. (2015). Don’t forget about the body: Exploring the curricular possibilities of embodied pedagogy. Innovative Higher Education, 40(4), 331-344. DOI 10.1007/s10755-015-9319-6
Paas, F., & Sweller, J. (2012). An evolutionary upgrade of cognitive load theory: Using the human motor system and collaboration to support the learning of complex cognitive tasks. Educational Psychology Review, 24(1), 27-45. DOI 10.1007/s10648-011-9179-2
Park, B., Korbach, A., Ginns, P., and Brünken, R. (2023). How learners use their hands for learning: an eye-tracking study. Educ. Psychol. Rev. 35:116. doi: 10.1007/s10648-023-09833-2
Pittalis, M., & Drijvers, P. (2023). Embodied instrumentation in a dynamic geometry environment: eleven-year-old students’ dragging schemes. Educational Studies in Mathematics, 113(2), 181-205. https://doi.org/10.1007/s10649-023-10222-3
Plummer, J. D., Lewis, H. K., Cho, K., Gil, M., Ricketts, A., Nolan, K., & Ruffner, M. B. (2026). Preschool-age children’s use of spatial sensemaking in science and engineering. Discover Education. https://doi.org/10.1007/s44217-026-01129-1
Post, L. S., Van Gog, T., Paas, F., and Zwaan, R. A. (2013). Efects of simultaneously observing and making gestures while studying grammar animations on cognitive load and learning. Comput. Human Behav. 29, 1450–1455. doi: 10.1016/j.chb.2013.01.005
Qi, Z., Wang, J., Du, Y., & Zhang, X. (2026). The effect of blended learning approaches on vocational school students’ practical skills performance and learning behaviors: virtual simulations or hands-on activities while accessing instructional videos. Instructional Science, 54(1), 39. https://doi.org/10.1007/s11251-026-09793-4
Rabattu, P. Y., Debarnot, U., and Hoyek, N. (2023). Exploring the impact of interactive movement-based anatomy learning in real classroom setting among kinesiology students. Anat. Sci. Educ. 16, 148–156. doi: 10.1002/ase.2172
Rahnert, K., Steenkamp, G., Jakobsson, N., and Ontong, J. (2024). the impact of observing lecturer hand motor actions on the learning of cognitive skills in higher education. Acta Psychol. 249:104483. doi: 10.1016/j.actpsy.2024.104483
Root-Bernstein, R. (2025). An art-science perspective on artificial intelligence creativity: From problem finding to materiality and embodied cognition. Journal of Creativity, 35(2), 100097. https://doi.org/10.1016/j.yjoc.2025.100097
Rouaud, J., Huet, N., & Przewozny, A. (2022). Assessing Spoken English Performance and Self-Efficacy Beliefs in the Classroom: Some Considerations on the Value of an Interdisciplinary Embodied Methodology for French Learners of English. Recherches anglaises et nord-américaines, (55), 87-113. https://doi.org/10.4000/ranam.433
Rudling, M., Rumetshofer, T., Nirme, J., & Mårtensson, J. (2026). Embodied word learning in schools and sustained attention in virtual reality. npj Science of Learning. https://doi.org/10.1038/s41539-025-00395-2
Sabouri-Moghadam, H., Nazari, M. A., Jahan, A., & Sabbaghipour, S. (2020). Barrasi-ye tasir-e harekat dar bazyabi-ye hafeze-ye hejāni: Motale'e-i tajrobi dar bab-e shenakht-e badanmand [Examining the effect of movement on emotional memory retrieval: An experimental study of embodied cognition]. Majalle-ye Ravan-shenasi va Ravanpezeshki-ye Shenakht, 7(3), 50–61. https://doi.org/10.52547/shenakht.7.3.50
Sadoski, M. (2015). Reading comprehension, embodied cognition, and dual coding theory. In S. R. Parris & K. Headley (Eds.), Comprehension instruction: research-based best practices (pp. 45–55). New York: Guilford Press.
Sadoski, M. (2018). Reading Comprehension is Embodied: Theoretical and Practical Considerations. Educ Psychol Rev 30, 331–349 https://doi.org/10.1007/s10648-017-9412-8
Sadoski, M., & Paivio, A. (2013). Imagery and text: a dual coding theory of reading and writing (2nd ed.). New York: Routledge.
Samuel, S., Salo, S., Ladvelin, T., Cole, G. G., & Eacott, M. J. (2023). Teleporting into walls? The irrelevance of the physical world in embodied perspective-taking. Psychonomic Bulletin & Review, 30(3), 1011-1019. https://doi.org/10.3758/s13423-022-02070-8
Samuelsson, R. (2026). Unpacking the child-technology-teacher relation: accentuating the role of embodied dynamics in young learners’ educational interactions. Discover Education, 5(1), 306. https://doi.org/10.1007/s44217-026-01481-2
Sandoval, P. G., & Mino, J. J. (2013). The Play's the Thing: Embodying Moments of Integration Live, on Stage. Learning Communities: Research & Practice, 1(2), 2.
Santos, M., Wolde, A., Taketomi, T., Yamamoto, G., Rodrigo, M., Sandor, C., & Kato, H. (2016). Augmented reality as multimedia: the case for situated vocabulary learning. Research and Practice in Techology Enhanced Learning, 11 (4), 1-23. DOI 10.1186/s41039-016-0028-2
Schaefer, S. (2025)Embodied Counting: Touching Objects Reduces Errors in Counting Under Cognitive Load. J Cogn Enhanc. 2025; 9: 206–13. https://doi.org/10.1007/s41465-025-00321-9
Schmidt, M., Benzing, V., Wallman-Jones, A., Mavilidi, M.-F., Lubans, D. R., and Paas, F. (2019). Embodied learning in the classroom: efects on primary school children’s attention and foreign language vocabulary learning. Psychol. Sport Exerc. 43, 45–54. doi: 10.1016/j.psychsport.2018.12.017
Shaghaghian, Z., Burte, H., Song, D., and Yan, W. (2024). An augmented reality application and experiment for understanding and learning spatial transformation matrices. Virtual Reality 28:12. doi: 10.1007/s10055-023-00904-x
Shapiro, L., & Stolz, S. A. (2019). Embodied cognition and its significance for education. Theory and Research in Education, 17(1), 19-39. doi.org/10.1177/1477878518822149
Skulmowski, A. (2019). Bridging the gap between embodied cognition and cognitive load theory (Doctoral dissertation, Dissertation, Chemnitz, Technische Universität Chemnitz, 2019).
Skulmowski, A., & Rey, G. D. (2018). Embodied learning: introducing a taxonomy based on bodily engagement and task integration. Cognitive research: principles and implications, 3(1), 6.
Skulmowski, A., Pradel, S., Kühnert, T., Brunnett, G., and Rey, G. D. (2016). Embodied learning using a tangible user interface: the efects of haptic perception and selective pointing on a spatial learning task. Comput. Educ. 92-93, 64–75. doi: 10.1016/j.compedu.2015.10.011
Smyth, L., Carter, J., Valter, K., and Webb, A. L. (2021). Examining the short-, medium-, and long-term success of an embodied learning activity in the study of hand anatomy for clinical application. Anat. Sci. Educ. 14, 201–209. doi: 10.1002/ase.1987
Spackman, J. S., & Yanchar, S. C. (2014). Embodied cognition, representationalism, and mechanism: a review and analysis. Journal for the Theory of Social Behaviour, 44(1), 46-79. DOI: 10.1111/jtsb.12028
Stalheim, O. R., & Somby, H. M. (2024). An embodied perspective on an augmented reality game in school: pupil's bodily experience toward learning. Smart Learning Environments, 11(1), 24. https://doi.org/10.1186/s40561-024-00308-7
Stockner, M., Mazzoni, G., Perrin, D., & Ianì, F. (2025). The same label, different processes: What lies behind the term “mental simulation” in the embodied cognition literature?. Brain and Cognition, 188, 106335. https://doi.org/10.1016/j.bandc.2025.106335
Swart, M. (2016). ¿ When is it the Gesture that Counts: Telling stories that cut to the [Cyber] chase–or, gest get to the po¡ nt!. Columbia University.
Swart, M. I., Kornkasem, S., & Colon-Acosta, N. (2017). From Abstract to Concrete-Evidence for designing learning platforms that adapt to user's. In Proceedings of the Annual Meeting of the Cognitive Science Society, 39 (0). Permalink.https://escholarship.org/uc/item/9p50w5vs
Sweller, J. (2023). The development of cognitive load theory: Replication crises and incorporation of other theories can lead to theory expansion. Educational Psychology Review, 35(4), 95. https://doi.org/10.1007/s10648-023-09817-2
Sweller, J., Van Merrienboer, J. J., & Paas, F. G. (1998). Cognitive architecture and instructional design. Educational psychology review, 10(3), 251-296. https://doi.org/10.1007/s10648-019-09465-5
Taheri, M. (2019). Era'e-ye olgu-ye khāndan-e zabān-e Fārsi bar asās-e ravesh-e āmuzesh-e badanmand barāye kudakān-e 4 ta 6 sāle [Developing a Persian reading model based on an embodied teaching method for children aged 4–6 years] [Master’s thesis, Islamic Azad University, Science and Research Branch and Shahrood Branch]. Iranian Research Institute for Information Science and Technology. https://ganj.irandoc.ac.ir
Trninic, D., & Abrahamson, D. (2012). Embodied interaction as designed mediation of conceptual performance. In Visual mathematics and cyberlearning (pp. 119-139). Dordrecht: Springer Netherlands. DOI 10.1007/978-94-007-2321-4 5
Tsaregorodtseva, O., & Kaup, B. (2024). Experiential traces first: Does holding a location in visuospatial working memory affect the processing of space-associated words?. Memory & Cognition, 52(4), 965-983. https://doi.org/10.3758/s13421-023-01512-5
Tu, X., Danish, J., Humburg, M., Zhou, M., Mathayas, N., Enyedy, N., & Jen, T. (2023). Understanding young children’s science learning through embodied communication within an MR environment: X. Tu et al. International Journal of Computer-Supported Collaborative Learning, 18(2), 231-258. 8. https://doi.org/10.1007/s11412-023-09395-z
Tuena, C., Di Lernia, D., Rodella, C., Bellinzona, F., Riva, G., Costello, M. C., & Repetto, C. (2023). The interaction between motor simulation and spatial perspective-taking in action language: a cross-cultural study. Memory & Cognition, 51(8), 1870-1880. https://doi.org/10.3758/s13421-023-01427-1
Vasilopoulos, F., Jeffrey, H., Wu, Y., & Dumontheil, I. (2023). Multi-level meta-analysis of physical activity interventions during childhood: Effects of physical activity on cognition and academic achievement. Educational psychology review, 35(2), 59. https://doi.org/10.1007/s10648-023-09760-2
Vergara, C., Urrutia, M., & Domínguez, A. (2025). When the Past Is Backward and the Future Is Forward: An Embodied Cognition Intervention in Preschoolers with Developmental Language Disorder. Journal of Intelligence, 13(6), 61. https://doi.org/10.3390/jintelligence13060061
Vitale, F., Padrón, I., Avenanti, A., & de Vega, M. (2021). Enhancing Motor Brain Activity Improves Memory for Action Language: A tDCS Study. Cerebral cortex (New York, N.Y. : 1991), 31(3), 1569–1581. https://doi.org/10.1093/cercor/bhaa309
Wang, J., & Liu, H. (2025). Curiosity-Driven Embodied Learning for Cross-modal Perception. Information Fusion, 103882. https://doi.org/10.1016/j.inffus.2025.103882
Wang, X., Fu, J., Luo, S., Zhao, K., & Duan, Q. (2026). XR-based cultural heritage information spaces: Towards a new paradigm of information encoding and decoding guided by embodied cognition. Information Processing & Management, 63(2), 104445. https://doi.org/10.1016/j.ipm.2025.104445
Wei, H., Bos, R., & Drijvers, P. (2024). Developing functional thinking: From concrete to abstract through an embodied design. Digital Experiences in Mathematics Education, 10(3), 323-351. https://doi.org/10.1007/s40751-024-00142-z
Wei, H., Bos, R., & Drijvers, P. (2024). Developing functional thinking: From concrete to abstract through an embodied design. Digital Experiences in Mathematics Education, 10, 323–351. https://doi.org/10.1007/s40751-024-00142-z
Wei, H., Bos, R., & Drijvers, P. (2026). An embodied approach to covariational reasoning: a hand-tracking study. Educational Studies in Mathematics, 121(1), 119-142. https://doi.org/10.1007/s10649-025-10435-8
Wingfield, C., & Connell, L. (2022). Sensorimotor distance: A fully grounded measure of semantic similarity for 800 million concept pairs. https://doi.org/10.3758/s13428-022-01965-7
Xu, X., & Ke, F. (2014). From psychomotor to ‘motorpsycho’: Learning through gestures with body sensory technologies. Educational Technology Research and Development, 62(6), 711-741. DOI 10.1007/s11423-014-9351-8
Xu, X., and Ke, F. (2020). Embodied interaction: learning Chinese characters through body movements. Lang. Learn. Technol. 24, 136–159.
Xu, X., and Ke, F. (2023). Learning number conversions through embodied interactions. Technol. Knowl. Learn. 28, 253–278. doi: 10.1007/s10758-021-09557-8
Yeo, L.-M., and Tzeng, Y.-T. (2020). Cognitive efect of tracing gesture in the learning from mathematics worked examples. Int. J. Sci. Math. Educ. 18, 733–751. doi:10.1007/s10763-019-09987-y
Yu, J. (2022). A Design Exploration of Leveraging Physical Play to Support Computational Learning for Young People (Doctoral dissertation, University of Colorado at Boulder). https://www.proquest.com/openview
Yu, J., Xu, T., Kelley, C., Ruppert, J., & Roque, R. (2025). Leveraging physical activities to support learning for young people via technologies: an examination of educational practices across the field. Review of Educational Research, 95(4), 775-821. https://doi.org/10.3102/00346543241248464
Yu, Q., and Yu, K. (2023). Does motion-sensing technology enhance students’ learning? Ameta-analysis. J. Educ. Comput. Res. 61, 1359–1388. doi: 10.1177/07356331231176741
Zhang, Y., Baills, F., and Prieto, P. (2024). Embodied music training can help improve speech imitation and pronunciation skills. Lang. Teach. 1-23, 1–23. doi: 10.1017/S0261444824000363
Zhong, B., Su, S., Liu, X., & Zhan, Z. (2023). A literature review on the empirical studies of technology-based embodied learning. Interactive learning environments, 31(8), 5180-5199. https://doi.org/10.1080/10494820.2021.1999274
Zhong, S., Tang, X., Cheng, X., & Pan, Y. (2026). Bodily maps of subject-specific feelings and academic emotions among high school students. BMC psychology, 14(1), 499. https://doi.org/10.1186/s40359-026-04283-1
Zhu, C., Klapwijk, R., Silva-Ordaz, M., Spandaw, J., & de Vries, M. J. (2023, December). Cognitive and embodied mapping of data: an examination of children’s spatial thinking in data physicalization. In Frontiers in Education (Vol. 8, p. 1308117). Frontiers Media SA. 7. doi: 10.3389/feduc.2023.1308117
Zuo, G., and Lin, L. (2025). Can whole-body tracing and hand tracing make any difference? Experimental evidence of learning outcomes, cognitive load, and intrinsic motivation on university students. Instr. Sci. 53, 1–25. doi: 10.1007/s11251- 024-09664-w.
